Nishino, T. & T. Morita. 1994. Biosystematic studies on the Kalimeris species (Compositae) in Niigata Prefecture. 1. Winter leaves and branching pattern. J. Jap. Bot. 69: 226–234.
Ito, M., A. Soejima & K. Watanabe. 1998. Phylogenetic relationships of Japanese aster (Asteraceae, Astereae) sensu lato based on chloroplast-DNA restriction site mutations. J. Pl. Res. 111: 217–223.
INOUE, S.,Y. SHINOHARA, S. TSUTSUMI, & K. KIDO. 1970. Cytological studies on the intergeneric natural hybrid between Aster and Kalimeris from Kyushu. (2) Natural hybrid between A. ageratoides subsp. ovatus and K. yomena. Mem. Fae. General Ed., Kumamoto Univ., (Ser. Nat. Sc.T No. 5: '3-80.
SHINDO, K. 1967. CytologicaL, morphological and geographical studies on differentiation of species in section Asteromoea of Kalimeris in Japan. Jour. Sci. Hiroshima Univ. Ser. B. Div. 2. 11: 127-199.
TARA, M. 1973. Cytogenetic studies on natural intergeneric hybridization in Aster alliances. II. Heteropappus hispidus (2n=36) X Kalimeris incisa (2n=72l. J. Sci. Hiroshima Univ. Ser. B, Div. 2, 14(3): 107~